The influence of movement and resting behavior on the range of three carabid beetles

We found that at large scales, in this case movement over hundreds of meters and several seasons, diffusion is a good approximation for population spread. At small scales of around one meter, BRW describes movement accurately. At intermediate scales we need both BRW, resting, and changes between these behaviors to describe motion effectively in the model. The area covered by the beetles increases linearly over time, while the maximum distance moved does not, indicating an approximately circular expansion and a thorough search of area traversed. Seasonal range and estimated rate of movement differed among species depending on prey availability but was not correlated to body size.